Title: ????? ?????? ??? ?????? Soybean ??????? ???????? Fabaceae ????? ?????? Glycine max, L. (Merr) ????? ????????? Soybean
1????? ????????? ?????? Soybean ???????
???????? Fabaceae ????? ?????? Glycine
max, L. (Merr) ????? ????????? Soybean
2- ????? ????? ??????
- ?? ???? ?????? ????? ????????? ???????? ????????
?????? ????? ??????? ???? ??????? ?? ????? ???
?????? ?? ?????? ??????? ???????? ???????? ??
????? ??????. - ?? ????? ?????? ??? ?????? ???????? ?????? ?????
??? ?????? ?? ??? ???????. - ?? ????? ?????? ??? ??????? ???????? ???????? ??
??? ?????? ??? ????? ????? ????? ???? ???? ??
???? ?? ?????? ????. - ?? ????? ?????? ??? ????? ????? ????????
??????????? ??????? ???????? ???? ?????? . - ?? ????? ?????? ??? ?????? ???????? ???????
???????? ?????? ???? ??????? ???????? ???? ??????
. - ?? ????? ?????? ??? ?????? ????? ??????? ?? ?????
?????? ????? ????? ?? . - ?? ????? ?????? ??? ????? ????? ??????? ???????
????? ??? ?????? ????? ??????? ???????? ???????
?????.
3?????? ???????? ?? ??? ???????
- ??? ?????? ?? ???????? ??????? ???? ??? ??? ???
?? ????????? ????? ??????? ???????? ???? ??? 1985
??? 119 ??? ???? ??????? ??? ??????? 1990 ??? 99
??? ???? ?????? ????? ?????? 1100 ??? ??????
Verdcout (1970) ?? ??? ?????? Glycine Soja and
zucc .?? ??? ????? ??? ?????? ?????? ????? ???
???? ??? ?????? ???????. - ???? ??????? ???????? ?? ??? ?????? ?? ?????? ???
1999 ??? 71850000 ????? ?????? ??????? ????? 2148
???/????? ????? ?????? ??????? ???????? ??
??????? ??? ??????? ?? 97000 ?????? ??? 1991 ???
17350 ?????? ??? ??? 1999 ????? ??????? ?????????
?? ??????? ??? ??????? ???? ?????? ??? 1112
???/???? ???? ????????? ????? ??????? ??? ??????
??? ?? ?? ??????? ???????? ?????????? ?????????.
4??????? ?????????? ?????? ??????
- ??? ??? ?????? ?????? ?? ????? ???????? ?
?????? ??????? ? ?????? ??????. - ??? ??? ?????? ?????? ????? ????? ???????
????????? ?? ??????? ???????? ?? ?????
????????????? ?? ?????? ?? ???????? ??????
40-50. - ???? ??? ?????? ????? ???? ??? ?????? ???
????????? ??????? ???? ??? ??? 50??? ????? ???
20 ??? ????? ??? ?????? ?? ????? ??????? ? ?????
? ???????? ?????? ????? 20 ?? ???? ????? .
5??? ??? ??????
- ????? ????? ?? ??? ??????
- ????? ??? ????? ????? Indeterminate .
- ?????? ????? ????? Determinate.
- ??? ??? ????? ???? ?????? ??????? ???????? ????
????? ???????? ??? ????? ?????? ???? ?????? ??
?????? . ??? ????? ??????? ???? ???????? ??? ??
???? ????????? ??? ?? ??? ???????? ??? ?? ????
????? ?? ??? ???????? (Bernard,1972) ?????? ??
??????? ??? ?????? ????? ???? ????? ?????? ?????
? ???????? ???? ???? ???????? ??????? ?? ?????
?????? ???? ?? ??????? ?????? ?????. ???????? ???
?????? ????? ????? ??????? ?? ????? ??????
??????? ????? ????? ??? ??? ?? ???? ?? ???????
????? ??? ????? ?? ??????? ? ??????? ????? ???
??? ??????? ?? ???? ????? ?????.
6??????? ??????? ????? ?? ??? ??????
- ?????? ??? ???????? ??
- 1- ?????? (OO) ??????? ?????? Norman, Geiso,
Shinsei . - 2- ?????? (O) ??????? ??????? Vansoy, Maple
Arrow, Elfe . - 3- ?????? (?) ??????? ??????? Anoka, Disoy, Hark
. - 4- ?????? (II) ??????? ??????? Magna, Harosoy,
Hawkeye . - 5- ?????? (III) ??????? ??????? Calland, Kin,
Ford ? ???? 35 - ????? 82 ????? 83.
7- 6- ?????? (IV) ??????? ???????Bonus, Clark, Kent
????? 22 ? ???? 111. - ?????? ????? Crawford ??????? ???????? ????? ???
55 ????? ????? ??? 120 ????? ????? ?????? 1200
??? ??????? . - 7- ?????? (V) ??????? ??????? Mack, York, Hill
??? ???? ????? ??? ???????? ????? ????????
Akishirome ????? ???? ??? ??????? ???????? ????
?? ?????? ???????? (???? ??? ????? 1992). - 8- ?????? (VI) ??????? ??????? Davis, Hood, Lee
. - 9- ?????? (VII) ??????? ??????? Ransom, Jakson,
Bragg . - 10- ?????? (VIII) ??????? ??????? Hampton,
Stuart, Hardee .
8??? ???????
- ????-???? 82
- ??? ?????? ???????? ???? ??? 95-100 ??? ??
??????? ??????? ????? ???? ????? ????????? ?????
??????? ???????? ??????? ??????? ?????? ?????
?????? ?? ??????? ????? ?????? ?? ????? ??? ?????
?????. - 2-???? 21
- ??? ????? ???????? ????? ??? ????? 120?????
?????? ?????? ?? ??????? ????? ????????? ?????
?????? ???? ??? ???? ??? ????? ????? ??????
?????? ????? ???????? ??????? ????? ??? ?????. - 3-???? 35
- ??? ?????? ???????? ????? ??? 105-110 ????? ??
??????? ?????? ?????? ?? ??????? ????? ?????????
????? ?????? ???? ??????? ?????? ??????? ??? ????
?????? ?????? ?? ??? ????. - 4--???? 22
- ??? ?????? ???????? ???????? ???? ???? ??? 115
????? ?????? ???????? ????? ??? ????? ????? ??
??????? ??? ?????? ??????? ??? ???? ?????? ??????
?? ??? ??? ????. - 5--???? 111
- ??? ?????? ???????? ?????? ????? ??? ????? ?????
??? 115-120 ????? ?? ??????? ????? ?????? ?? ????
????????? ??? ?????? ?????? ??? ???? ??????
?????? ?? ??? ??? ????. - 6--???? 83
- ??? ?????? ???????? ???? ????? ???? ???
95-100??? ??????? ????? ?????? ????? ?????????
??? ??? ????? ????? ??? ????? ??? ???? ??????? ??
????? ?????? ????? ??????? ?? ?????? ????? ??
?????? ?? ????? ??? ???? ??? ?????.
9????? ?????
- ????? ????? ???????
- ????? ??? ??????? ??? ????? 15 ????? ???? 10
???? ??? ????? ??????? ?5 ???? ??? ??? ?????
??????? ???????. - ?????? ????? ????? ???????
- ??? ?????? ????? ?? ???? ??? ????? ????????
????????. ???? ??????? ??? ??????? ????? ???????
??????? ?????? ??? ??? ?????? ??????? ?????????
??? ????? ?? ????? ??? ??? ??????? ??????
????????.
10????? ????? ??? ??????
11- ????? ??????? ?? ??? ??????
12- ???? ????? ????? ???? ??????
13- ???? ????? ??? ??????? ???
- 1- VS1 ??? ??????? ???? ????? ???? ?????? ???
??????? ??????? ?? ???? ??? ??????? ???????
?????? ???? ??? ?????. - 2-. VS2 ????? ????? ???? ?????? ??? ?????? ???
(3) ??? ????? ??????. -
- 3. Vs3 ????? ????? ???? ??? ??? ????? ??????? ???
????? ?????? ????????. -
- 4. VS4 ????? ????? ???? ??? ??? ????? ??????? ???
????? ?????? ????????. -
- 5. VSn ?????? ??? ??? ?? ????? ný ??? ????? ???
????? ?????? ???????? ???? ???? ???????? ??? ???
????? ??????? ???? ??? ????? ????? ??? ????? ??
??????? ??? ?????? ????? ???? ?? ??????? ??????
?????.
14????? ????? ???????
15????? ????? ???????
16???? ????? ????? ???????
17- ?????? ????? ???????????? ???? ??????? ???
???? ????? ????? ??????? ???? ????? ?????? ??????
??? ????? ??????? . - 1. RS1 ????? ??? ??? ??????? ????? ???????
????? ??????? ????? ????? ??? ?? ???? ?????
???????. - 2. RS2 ????? ????? ??? ????? ??????? ????? ?????
??????? ?????? ???? ?????? ??? ???? ?? ????????
????????? ????? ??????? ?? ??? ???? ??????? ??
??????? ??? ?????? ?????. - 3. RS3 ????? ??? ???? ?????? ???????? ???? ???
??????? ??? ??? ?????? ??? ??????? ????? ???????
?? ?????? ??? ???????. - 4. RS4 ????? ???? ??? ????? ????????? ??? ???
?????? ??? ??????? ????? ??????? ?? ?????? ???
???????.
18- RS .5 ??? ??? ??????? ???? ????? ????? ??????
????? ????? ????? 1/8 ???? ?? 3 ?? ?? ??? ??????
???? ?????? ??? ??? ?????? ??? ??????? ?????
??????? ?? ?????? ??? ???????. - 6. RS6 ????? ???? ?????? ??????? ?? ?????? ????
?????? ?? ????? ???? ??? ??? ?????? ??? ???????
????? ??????? ????? ????? ??????? ?? ?????? ???
???????. - 7. RS7 ????? ????? ??? ????? ????? ????? ????? ??
??? ??????? ?? ????? ??? ???? ??? ????? ???????
??? ??? ??? ????? ?????? ?? ???? ???? ??????
?????. - 8. RS8 ????? ????? ??? ????? ????? ????? ?????
???? 25 ?? ?????? ?? ???? ??? ??? ?????? ????
????? (????? ?????).
19- ???? ????? ????? ???? ??????
20Table 1. Vegetative and reproductive stages of
a soybean plant.
21Germination and Emergence
Fig.3
Fig.1
22 V2 Stage (second node)
Fig.4
Fig.1
Fig.6
Fig.5
23 V3 and V5 Stages (third and fifth
nodes)
Fig.7
Fig.1
Fig.21
Fig.8
24V6 Stage (sixth node)
Fig.9
25R1 Stage (beginning bloom)
Fig.12
Fig.21
26R4 Stage (full pod)
Fig. 25
Fig.20
Fig.17
Fig.27
27R6 Stage (full seed)
Fig. 31
Fig. 27
Fig.28
Fig. 38
Fig. 39
Fig. 40
28R7 Stage (beginning maturity
Fig.33
Fig. 32
Fig.34
Fig.35
29R8 Stage (full maturity)
Fig.37
Fig.36
30?????? ????????
- ???? ??? ?????? ?? ??????? ?????? ???? ????? .
- ????? ??????? ???????? ???????.
- ???? ?????? ?? ??????? ??????? ???????? .
- ?? ???? ?????? ?? ??????? ??????? ??????? ?????
. - ??? ????? ?????? ???????? ????? ??????? ????????
??? ??????? ??????.
31????? ??????? ??????? ????????
- ??? ?????? ????? ???? ???? ?????? ??????? ?? 15
???? ??? ????? ?????. ?????? ??????? ???????
??????? ???? ??? ????. - ???? ?? ??? ??? ????? ??????? ?????? ?????? ??
?????? ????????.???? ????? ???? ????? ?????? 1985
??? ????? ????? ????? ??????? ??? ?????? ??? ???
?????? ??? ???? ?????? ???????? ? ????? ??????
??? ???????? ??????? ??????? ????? ?????? ??????
?????? ?????? ???? ??? ??????? ??????? ????????
???????? ??????? ?????????? ? ????? ????? ???
???????? ??????? ?????? ?????? ??????? ??????
??????? ???? ??? ????? ?????.
32????? ????? ??????? ????????
33??????? ??? ?????? ??????? ????????
34????? ????? ??????? ????????
35????? ????? ??????? ?????? ?????? ?? ????? ????
?????????
36???? ???????
- ????? ?????? ??? 35 ??? ????? ?????? ??????? ??
????? ??????? ?? ??????? ????21-????22-????111
?40 ??? ????? ?????? ?? ??????? ???? 82 ?????83
?????35 ???? ????? ?????? ??????? ????? ???????
??? ???? ??? ????? ??????? ???? ?????? ????????
??? ????? ????? ????? ?? ??????? ???????? ??? ???
60??? ????/???? ??? ????? ?? ?????? ??? 25 ???
????/???? ??????? ?????? ???? ?????? ?????
??????? ??? ??????? ?????? ???? ???? ?????? ???
??????? ??? ???? ??????? ????.
37????? ???? ??????? ????????
38???? ??????? ???????? ????????
39?????????? ???????
- 1- ???? ???????
- ????? ???? ??????? ??? ?????? ??? ?????? ????
?????? ???? ??????? ?? ?????? ?? ??? ????????
?????? ????? ????? ?????? ???? ??????? ??????. ?
?????? ???? ??????? 20- 530? ?? ?????? ???????
??????? ?? ?????? ???? ??????? ?? ??? ??? ????
???????. ????? ???????? ????? ?? ????? ???????
???? ????? ?????? ??? ?????? ???? ??????? ??
540?. ??????? ???? ??????? ?? 535? ????? ???
?????? ???? ?? ?????? ??????? ???????? ?????? ???
????? ?????? .(Pandey, 1987) - ??? ??? ?????(Hofstea, 1984) ??? ?????? ?? ????
????? ?????? ?????? ?? 15-510? ??? 21-516? ?????
??? ?????? ???? ?? ?????? ?????? ??????? ??????
?????? ?? ?????? ?????? ??? ????? ??????. - 2- ?????
- ?. ??? ?????? ??????? ???? ???? ??? ??????
?????? ?????? ?????? ?? ??? ???? ??? ???????
?????? ???? ?????? ?????? ??????. - ?. ??? ??????? ???? ??? ??????? ???? ??????
??????? ??? ??? ?????? ???? ??? ?????? ???? ???
??? ????? ?????? ???????.
40?????????? ????????
- ???? ??????? ???? ????? ?????? .
- ????? ??????? ???????? ???? ?????? ??? ?????
?????? ????? ???? ??????? ?? ??????? ??????? ???
?????? ??? ????? ??????? ??? ??????. - ???? 30 ??? ????? ??? ??????? ?75 ??? ???? ??????
????????? ?25 ??? ?? ????? ?????????? (Panday,
1987) ? ????? ????? ??? ???????? - ???? ??????? -
????? ???????? ??? ?? ??????? ???????? ?????????
???????? 60 ??? ???? 24 ??? ??? ?????????? 36
??? ????? ???????? ??? ??? ??? ????? ??? ??????
??????? ??????? ??? ???? ????? ???????? ??????
???????? ?????? ??????? ?????? (????? 1993). - ?? ???? ????? ??? ?????? ???????? ????? ?????????
(??????? ???????)??? ???? ???? ????? ??? ???????
?????? ??? ?????? ???? ??? ?????? ??? ??????
?????? ?? ????? ??????? ????????? ????? 406040
??/???? ?? ?????? ???????? ?? ????? 3?? ??? ???
??? ?? ???? ??????? ??????? ??? ??????? ????
?????? ?????? ??? ??? ??????? ???????? ???
??????? ??? ?? ???? ???? ??? ?? ?????? ?????? ??
???? ??? ??????.
41(No Transcript)
42???? ???? ?? ?????? ?????????
43??? ???????
- ???? ??? ?????? ????????
- ?????? ?????? ???? ?????? ??? ????? ????? ???
????? ??? ????? ???? ?????? ??????? ??? ???????
??? ?????? ?? ??? ??????? 15 ?? ?? ???? ???????
10 ??/?????? ?? ??? ????? ?? ???? ??????? ?????
12 ??/?????? ?? ??? 3-4 ???? ?? ?????? ?? ????
???? ?????? ???? ??????? ?? ????? ?????? ?? ????
?? ???? ????? ????? ??? ??????? ??????. - ??????? (???????) ???? ????? ???? ?????? ??????
??? ??? ?????? ??????? ???? ???? ??????? ???? ??
??????? ? ???? ??????? ??? ?? ??? ?? ??? ?????
?????? ?????? ??????? ?? ???? ?? ????? ??????
???????? ??????? ?????? ?? ????? ?????? ???? ????
????? ?????? ??? ?????? ???? ?? ???? ??? ???????
3?? ?? ?????? ? 5 ?? ?? ???????. - ???? ????? ???????? ??????? ??? ????????? ?????
?????? ?? ???????? ?? ???? ???????(140-150 ???
????/????) - ???? ????? ????? ??????? ???????? ?????? ????????
??????? ???????? ????????? ??? ??????? ???????
?????? (???????) ??? ??? ??????? ??? ??????
???????? ?? ????? ??????? ????? ????? ??????
????? ???? ????? ??????? ???? ?????? ?? ????
?????? ??????? ???? ?????? ??? ??????? ???????
???? ??????? ???? ???????? ?????? ???????
???????? ??????? ??????? ??????? ??????? ??? ????
?????? ?????
44????? ????? ????? ???????
45?????? ??? ?????? ??? ???????
46?????? ??? ??????
47?????????? ???????
- ????? ?????????? ??????? ????? ???????
??????? ???????? ????? ??? ???? ????? ?????? ???
????? ??????? ??? 15-20 ????? ?? ??????? ?? ????
?? 12-15 ??? ???? ?? ??????? ??????? ? ???? 5-6
??? ???????? ??????? ?????? ??????? ??????
??????? ??? 5-6 ???? ?? ??????? ???? ??? ?????
???????? ?? ???? ???? ??? ????? ???? ????? ?????
??????? ??????.
48?????? ???????
- ??????? ??? ???? ????? 3-4 ???? ?????? ???????
?????? ?????? ??? ????? ???????? ? ????? ???????
????? ????? ??????? ????? ?????? ?????? ???????
??? ????? ??? ???? ??? ???????? ???????? ??? ????
???? ??????? ???? ????? ???? ???????? ??? ???
????? ??? ?????? ??????? ??? ??????? ?? ????????
?????? ??????? ?????? ??? ???? ???? ??? ???
?????? ????? ?? ??????? ???? ???? ?? 45??? ??????
?? ??? ???????? ?????? ??? ????? ???? ??? ?????
?????? - -??????? ??????? ??????? ?????? ???? ?????
500????? 1,7??? /???? ?? ??? 48 ?? ???????48
????? 205 ??? / ???? ??? ??????? ???? ???? ??
??????? ?????? ?? ??? ??? ??? ??????? ?? ???????
??????? ???? ?? 200-300??? ???/????. - ??? ???? ???? ?????? ? ?????? ?????? ??? ???????
50 ?? ????? 24 ????? ???/ ???? ?? 300 ??? ???
?? ??? ???????? .
49?????? ??????? ???????? ???????
- ????? ??????? ????????
- ???? ??? ?????? ?????? ???? ????? ????? ???????
??????? ???????? ??????? ????????? ???? ???????
?????? ??????? ??????? ??????? ?? 1??/1??? ???
???????? ???????? ??????? ????? ??????. - ?????? ?????? ???????
- ???? ??? ?????? ???????? ???????
- 1. ???? ??? ????? ?????? ???? ????? ????? ????
??????? ????????? ??? ????????50 ????? ??? ????
??????. - 2. ???? ?????? ????? ?????? ????? ?????????.
- 3. ???????? ?????? ??????? ????????? ??
???????????????? ?? ??????? ?? ????? ????????
?????? ?????. - 4. ?????? ??????? ?????? ??? ?????? ????????
????? ????? ????? ????? ?? ????? ?? ???? ????? ??
????? ????? ???????? ??? ????????? ?? ??????? ??
???????? ?? ?????????? ?? ??? ?.?.?. -
50?????? ?????? ???????
- ?? ?????? ????? ?????? ??????? ??????? ??????
?????? ?????? ???????? ??? ??? ??????. ???? ???
????? ????? ?????? ??? ?? ??? ?????? ????? ??????
???. ??? ??? ??? ??????? ?? ?????? ??? ??????
???? ?? ??? ?????? ?? ?????? ??????????. - ?? ???? ??????? ?????? ?? ??? ??? ???? ??? ?????
???? ??????? ??????? ???? ??????? ? ??? ?????
?????? ????? ??? ?????? ?????? ?????? ????????
??? ??? ?? ???????. ???? ??? ??????? ??? ?????
?????? ?? ??? ?????? ??? ??? ??? ?? ??? ?????
??????? ?? ??????? ?? ????? ?? ???? ??? ????
?????? ??????? ?????? ??? ???????? ????????.
51????? ????? ???? ??????
52?????? ????? ???? ??????
53???? ??? ????? ??? ?????? ?????
54???????
- ???? ????? ??????? ?? 800-1100 ??? ????/????. ???
???? ??? ?????? ???? ???? ???? ??????? ?? ????
??????? ?? ????? ??????? ?? ????? ???????.
55????? ???????
- ?-??? ?? ??????? ??????
- ?1 ???? ?? ??? ??? ???????
- ?2 ???? ??? ??? ?????? ????? ???? ????? ??????
- ?3 ????? ?????? ??????? ???????? ?? ??? ??????
??? 1990 ???? ????? - ?4 ?? ?? ??????? ?????????? ???? ?????? ?? ????
?????? ???????? ????? - ?5 ????? ???? ????? ???? ????? ??? ??? ???
??????? - ?6 ??? ?????? ????? ?? ??? ?????? ??? ?? ??? ????
- ?7 ???? ??????? ??????????? ???? ??? ???????
- ?8 ?? ?? ??????? ??? ???? ??????? ?????? ?? ???
??????? - ?9 ?? ?? ?????????? ??????? ???????? ???? ???
??????? - ?10 ?? ?? ???? ????? ?????? ??? ???????
- ?11 ???? ???????? ???????? ?????? ??? ????
??????? ?? ???? ??????? ??? ????? ????? ???
?????? ??????? ????????? - ?12 ???? ?? ???? ???????? ???? ?????? ?
56- ?- ?? ????? ?? ?????? ??? ??? ???????? ???????.
- 1-( )???? ???? ??? ?????? ?????? ???? ?? ??????
????? ?? ???? ??????? ???????? ???? ????. - 2-( )??????? ?? ??? ?????? ???? ??? ?? ??????
?????. - 3-( )?????? ?? ???? ??? ?????? ????? ????? ?????
?? ????? ???????? ?????? ??????. - 4-( ) ???? ??????? ??? ??? ??? ?????? ????? ?????
??????? ???? ??????? ??????? ??????? . - 5-( ) ????? ??? ?????? ????? ????? ???? ????
????? ?????? ????? ?? ??? ?????? ??? ????? ?????.
57- ??-???? ??????? ??????? ?? ??? ???????
- 1-???? ??? ?????? ???? ????? (???? ?????-?????
??????-?????). - 2-?????? ?? ??? ?????? (??????-???-??????) ?????
??? ?????? ?? ??????. - 3-???? ????? ?? ??? ?????? ?? (??????-???????-????
??). - 4-???? ???? (?????-?????-???????) ??????? ??? ???
??? ??????. - 5-???? ????? ??? ?????? ?? ??????? ???????
(???????-???????-???????)