Title: The Diversity of Life
1The Diversity of Life I. An Overview
2The Diversity of Life I. An Overview A.
Classifying Organisms
3The Diversity of Life I. An Overview A.
Classifying Organisms Initially, using a
Platonic, typological concept, Linnaeus and
others created a nested, hierarchical system.
4The Diversity of Life I. An Overview A.
Classifying Organisms Initially, using a
Platonic, typological concept, Linnaeus and
others created a nested, hierarchical
system. Evolution explained this nested pattern
as a consequence of descent from common
ancestors.
5The Diversity of Life I. An Overview A.
Classifying Organisms Initially, using a
Platonic, typological concept, Linnaeus and
others created a nested, hierarchical
system. Evolution explained this nested pattern
as a consequence of descent from common
ancestors. Modern biologists view the
classification system as a means of showing the
phylogenetic relationships among groups.
6The Diversity of Life I. An Overview A.
Classifying Organisms Initially, using a
Platonic, typological concept, Linnaeus and
others created a nested, hierarchical
system. Evolution explained this nested pattern
as a consequence of descent from common
ancestors. Modern biologists view the
classification system as a means of showing the
phylogenetic relationships among groups. Genetic
relatedness should be the basic for biological
classification...
Genus Felis
Genus Panthera
Family Felidae
7The Diversity of Life I. An Overview A.
Classifying Organisms B. Kingdoms
8The Diversity of Life I. An Overview A.
Classifying Organisms B. Kingdoms
Phylogenetic analysis revealed that the "Monera"
were an incredibly diverse group genetically.
Also, one subgroup - the Archea, were more
similar to Eukaryotes than to the other group of
prokaryotes (the 'Eubacteria').
9The Diversity of Life I. An Overview A.
Classifying Organisms B. Kingdoms
The most common way of ordering the major groups
of life has been the 5 kingdom approach.
10The Diversity of Life I. An Overview A.
Classifying Organisms B. Kingdoms
Phylogenetic analysis revealed that the "Monera"
were an incredibly diverse group genetically.
Also, one subgroup - the Archea, were more
similar to Eukaryotes than to the other group of
prokaryotes (the 'Eubacteria'). This required a
new way of looking at the most fundamental
groupings of life - and the introduction of a new
term Domains
11The Diversity of Life I. An Overview A.
Classifying B. Kingdoms C. Domains
12The Diversity of Life I. An Overview A.
Classifying B. Kingdoms C. Domains
Curiously, the very root of life may be invisible
to genetic analysis. Bacteria transfer genes by
division (to 'offspring'), but they also transfer
genes "laterally" to other living bacteria. This
makes reconstructing bacterial phylogenies
difficult.
13The Diversity of Life I. An Overview A.
Classifying B. Kingdoms C. Domains
Also, early evolution involved bacterial
symbioses and gene sharing between hosts and
symbionts
14The Diversity of Life I. An Overview A.
Classifying B. Kingdoms C. Domains
So, reconstructing the patterns of relatedness
among these ancient life forms is difficult.
15The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
16The Diversity of Life II. An Overview of the
Bacteria
5 million to present
2.3-2.0 bya Oxygen
1.8 bya first eukaryote
0.9 bya first animals
0.5 bya Cambrian
0.24 byaMesozoic
0.065 byaCenozoic
4.5 bya Earth Forms
4.0 bya Oldest Rocks
3.4 bya Oldest Fossils
for 1/2 of life's history, life was exclusively
bacterial.... what were they doing? Spheres,
rods, and spirals were all they could come up
with?? Let's look...
17The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
18The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of organisms.
19The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. A. Oxygen Demand all eukaryotes
require oxygen.
20The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. A. Oxygen Demand all eukaryotes
require oxygen. bacteria show greater
variability - obligate anaerobes - die in
presence of O2 - aerotolerant - don't die,
but don't use O2 - facultative aerobes - can
use O2, but don't need it - obligate aerobes
- require O2 to live
21The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. A. Oxygen Demand all eukaryotes
require oxygen. bacteria show greater
variability - obligate anaerobes - die in
presence of O2 - aerotolerant - don't die,
but don't use O2 - facultative aerobes - can
use O2, but don't need it - obligate aerobes
- require O2 to live
represents an interesting continuum, perhaps
correlating with the production of O2 in the
atmosphere.
22The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. B. Nutritional Categories
23The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. B. Nutritional Categories -
chemolithotrophs use inorganics (H2S, etc.) as
electron donors for electron transport chains and
use energy to fix carbon dioxide. Only done by
bacteria.
24The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. B. Nutritional Categories -
chemolithotrophs use inorganics (H2S, etc.) as
electron donors for electron transport chains and
use energy to fix carbon dioxide. Only done by
bacteria. - photoheterotrophs use light as
source of energy, but harvest organics from
environment. Only done by bacteria.
25The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. B. Nutritional Categories -
chemolithotrophs use inorganics (H2S, etc.) as
electron donors for electron transport chains and
use energy to fix carbon dioxide. Only done by
bacteria. - photoheterotrophs use light as
source of energy, but harvest organics from
environment. Only done by bacteria. -
photoautotrophs use light as source of energy,
and use this energy to fix carbon dioxide.
bacteria and some eukaryotes.
26The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. B. Nutritional Categories -
chemolithotrophs use inorganics (H2S, etc.) as
electron donors for electron transport chains and
use energy to fix carbon dioxide. Only done by
bacteria. - photoheterotrophs use light as
source of energy, but harvest organics from
environment. Only done by bacteria. -
photoautotrophs use light as source of energy,
and use this energy to fix carbon dioxide.
bacteria and some eukaryotes. -
chemoheterotrophs get energy and carbon from
organics they consume. bacteria and some
eukaryotes.
27The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. C. Their Ecological Importance
28The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. C. Their Ecological Importance -
major photosynthetic contributors (with protists
and plants)
29The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. C. Their Ecological Importance -
major photosynthetic contributors (with protists
and plants) - the only organisms that fix
nitrogen into biologically useful forms that can
be absorbed by plants.
30The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. C. Their Ecological Importance -
major photosynthetic contributors (with protists
and plants) - the only organisms that fix
nitrogen into biologically useful forms (that can
be absorbed by plants). Why is Nitrogen
important? - primary decomposers (with fungi)
31The Diversity of Life I. An Overview II. An
Overview of 'The Bacteria'
The key thing about bacteria is their metabolic
diversity. Although they didn't radiate much
morphologically (spheres, rod, spirals), they DID
radiate metabolically. As a group, they are the
most metabolically diverse group of
organisms. C. Their Ecological Importance -
major photosynthetic contributors (with protists
and plants) - the only organisms that fix
nitrogen into biologically useful forms that can
be absorbed by plants. - primary decomposers
(with fungi) - pathogens (cause disease)
32The Diversity of Life III. Domain Eukarya
Protists are single celled or colonial organisms
they are the most primitive eukaryotes, and they
probably evolved by endosymbiotic interactions
among different types of bacteria
33The Diversity of Life III. Domain Eukarya
From different types of protists evolved
different types of multicellular eukaryotes the
fungi, plants, and animals.
34The Diversity of Life III. Domain Eukarya
D. Diversity - green alga
Same chlorophyll as plants alternation of
generation genetic analysis confirms relatedness
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36The Diversity of Life IV. Domain Eukarya
D. Diversity - Choanoflagellates
37II. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce)
38IV. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce)
39IV. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce) 2. Colonization of Land Environmental
Diffs Aquatic Habitats Terrestrial Water
available Desiccating
40IV. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce) 2. Colonization of Land Environmental
Diffs Aquatic Habitats Terrestrial Water
available Desiccating Sunlight
absorbed Sunlight available
41IV. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce) 2. Colonization of Land Environmental
Diffs Aquatic Habitats Terrestrial Water
available Desiccating Sunlight
absorbed Sunlight available Nutrients at
Depth Nutrients available
42IV. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce) 2. Colonization of Land Environmental
Diffs Aquatic Habitats Terrestrial Water
available Desiccating Sunlight
absorbed Sunlight available Nutrients at
Depth Nutrients available Buoyant Less
Supportive
43IV. Introduction to Plants A. Evolutionary
History 1. Green Algal roots Ulva (sea
lettuce) 2. Colonization of Land Environmental
Diffs Aquatic Habitats Terrestrial Water
available Desiccating Sunlight
absorbed Sunlight available Nutrients at
Depth Nutrients available Buoyant Less
Supportive Low oxygen, higher CO2 reverse
44IV. Introduction to Plants A. Evolutionary
History B. Adaptations to Life on Land C.
Plant Evolution Acquisition of Terrestriality
45V. Plant Diversity A. Non-Vascular (no true
xylem or phloem) 2. Example Mosses
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47V. Plant Diversity B. Non-seed
Tracheophytes 1. Lycophytes (Club Mosses)
48III. Plant Diversity B. Non-seed
Tracheophytes 1. Lycophytes - (Club Mosses) -
ancient dominated first forests 300-350
mya
49III. Plant Diversity B. Non-seed
Tracheophytes 2. Ferns and their allies
50III. Plant Diversity B. Non-seed
Tracheophytes 2. Ferns and their allies -
true complex leaves - true roots - also
ancient appearing 350 mya - dominant
sporophyte reduced gametophyte
51Fern Life Cycle
52III. Plant Diversity C. Seed Tracheophytes 1.
Gymnosperms naked seed a. Evolutionary
History - dominated during Permian (280
mya) and through Mesozoic, and still
dominate in dry env. Today (high latitudes,
sandy soils)
53 54 III. Plant Diversity C. Seed Tracheophytes 2.
Angiosperms flowering plants - flowers -
bribe pollinators - fruits - bribe seed
dispersers
55