Title: Community Ecology
1Community Ecology
- Divya Patel, Nastassja Kosinski, Amanda Manzione,
Renée Buttel
2Concept 53.1- A community's interactions include
competition,predation, herbivory, symbiosis, and
disease.
Competition (-/-) Competition for resources between plants, between animals, or between decomposers when resources are in short supply. An ecological niche is the sum total of a species use of the biotic and abiotic resources in its environment. The competitive exclusion principle states that two species cannot coexist in the same community if their niches are identical. Character Displacement is the tendency for characteristics to be more divergent in sympatric (geographically overlapping) populations of two species than in allopatric (geographically separate) populations of the same two species.
Predation (/-) An interaction between species in which one species, the predator, eats the other, the prey. Because of predation, many species have developed special adaptations, such as various forms of mimicry and coloration.
Herbivory (/-) An interaction in which a herbivore eats a part of a plant or an alga. Because of herbivory many plants and algae have developed many chemical toxins and physical defenses (spines and thorns), while many herbivores have developed the ability to distinguish the difference between nutritious and malnutritious plants.
Parasitism (/-) A type of symbiotic relationship in which the symbiont (parasite) benefits at the expense of the host by living either within the host (endoparasitism) or outside the host (exoparasitism). Parasitoidism is a type of parasitism in which an insect lays eggs on or in a living host the larvae then feed on the body of the host eventually killing it.
Disease (/-) Pathogens (disease- causing agents) can be bacteria, viruses, or protists, but fungi and prions can be pathogenic also. Pathogens inflict lethal harm on their hosts. Pathogens are very much like parasites.
Mutualism (/) A symbiotic relationship in which both participants benefit. Mutualistic relationships sometimes involve the evolution of related adaptations in both species, with changes in either species likely to affect the survival and reproduction of the other.
Commensalism (/0) An interaction between two species in which one of the benefits, but the neither harms nor helps the other. There few if any cases of pure commensalism.
- Interspecific Interactions and Adaptation-
Evidence for coevolution, involving reciprocal
genetic change by interaction populations, is
scarce. However, generalized adaptations of
organisms to other organisms in their environment
is a fundamental feature of life. - Populations are linked by interspecific
interactions that affect the survival and
reproduction of the species engaged in the
interaction.
3Concept 53.2- Dominant and keystone species
exert strong controls on community structure
- Species Diversity- Species diversity measures the
number of species in a community- its species
richness- and their relative abundance.Species
richness is the total number of different species
in the community. Relative abundance is the
proportion each species represents of the total
individuals in the community. Measuring species
diversity is easier said then done, there are
very few methods used to accurately measure
species diversity. Measuring species diversity is
important not only for understanding community
structure but also to conserve biodiversity. - Trophic Structure- The structure and dynamics of
a community depend to a large extent on the
feeding relationships between the organisms- the
trophic structure of the community. A food chain
is the transfer of food energy up the trophic
levels from its source in plants (primary
producers) to primary consumers to secondary and
tertiary consumers. A food web is the connection
of many food chains it is the elaborate,
interconnected feeding relationships in an
ecosystem. The energetic hypothesis suggests that
the length of a food chain is limited by the
inefficiency of energy transfer along the chain.
The dynamic stability hypothesis proposes that
long food chains are less stable than short
chains. - Species with a Large Impact Dominant species
and keystone species exert strong controls on
community structure. Dominant species are the
most abundant species in a community and their
dominance is achieved by having high competitive
ability. Keystone species are usually less
abundant species that exert a disproportionate
influence on community structure because of their
ecological niche. Ecosystem engineers, also
called foundation species, exert influence on
community structure through their effects on the
physical environment. - Bottom-Up and Top-Down Controls- The bottom-up
model proposed an unidirectional influence from
lower to higher trophic level, in which nutrients
and other abiotic factors are the main
determinants of community structure, including
the abundance of primary producers. The top-down
model proposes that control of each trophic level
comes from the trophic level above, with the
result that predators control herbivores, which
in turn, control primary producers.
4Concept 53.3- Disturbance influences species
diversity and composition
- Disturbance- Increasingly, evidence suggests that
disturbance and nonequilibrium rather than
stability and equilibrium are the norm for most
communities. According to the intermediate
disturbance hypothesis moderate level of
disturbance can foster higher species diversity
than can low or high levels of disturbance. - Human Disturbance- Humans are the most widespread
agents of disturbance, and their disturbance to
communities usually reduces species diversity.
Humans also prevent some naturally occuring
disturbances, such as fire, which can be
important to community structure. - Ecological Succession- Ecological succession is
the sequence of community and ecosystem changes
after a disturbance. Primary succession occurs
where no soil exists when succession begins
secondary succession begins in a are where soil
remains after a disturbance. Mechanisms producing
community change during succession include
facilitation and inhibition.
5Concept 53.4- Biogeographic factors that affect
community diversity
- Research done by Charles Darwin and Alfred
Wallace proved that tropical habitats support
many more species than temperate and polar
regions. An example of this is that there are 711
different types of tree species in Malaysia while
there are only 50 tree species in the area north
of the Alps in Western Europe. - Species richness in equatorial-polar gradients
are affected by evolutionary history and time. - As time passes, more speciation events will
occur, causing species diversity to increase. - Tropical habitats are older than both polar and
temperate regions, proving that species richness
is more abundant in the tropics than at an
equatorial-pole gradient. - Biodiversity is affected by climate. Solar energy
and water are two components of climate that
effect biodiversity. - The species area curve was recognized by
Alexander von Humboldt in 1807. This curve states
that the larger area, the greater the number of
species. - The amount of species living on an island is
based on the rate of immigration and the rate of
extinction. These rates are affected by the size
of the Island, the amount of species living on
the island, and the distance from the island to
the mainland. Larger islands have lower
extinction rates because they contain more
resources and more diverse habitats. - The island equilibrium model was creates by
MacArthur and Wilson. This model shows that when
the rate of immigration is equal to the rate of
extinction, an equilibrium will be reached. The
islands size and distance from its mainland
effect species richness( at equilibrium). Many
scientist have questioned the accuracy of this
model.
6Concept 53.5- Contrasting views of community
structure are the subject of continuing debate
- The integrated hypothesis was created by F.E.
Clemens, of the Carnegie Institute of Washington,
in the early 1900s. This hypothesis states that
a community acts as a superorganism, as a result
of required biotic interactions . - Then, H.A. Gleason, of the University of Chicago,
created the individualistic hypothesis. This
hypothesis states that species with the same
abiotic needs live in the same area. - These hypotheses are similar because they state
that communities interact. - These hypotheses differ because the integrated
hypothesis states that species depend upon their
interactions with other species and he
individualistic hypothesis states that each
species lives in an environment where the abiotic
factors that it needs are present. - American scientists Paul and Anne Ehrlich
proposed the idea that all species in a community
depend upon and affect one another. - The rivet model proposed the idea that species in
a community interact with species in the web of
life. Increasing or decreasing the population of
one species will greatly affect the population of
another species. - The Redundancy model was created in 1992 by Brian
Walker, an Australian Ecologist. This model
proposed the idea that most species are not
associated with one another and that decreasing
or increasing the population of one species will
not greatly affect the population of another
species.
7What is DDT?
- DDT is a pesticide that was released to the
public after WWII. - Unfortunately, during that time and even today,
many people are unaware of the side effects of
DDT in the environment especially on factors such
as herbivory, predation, and interspecific
competition. - Thanks to the work of many researchers and
publicists, such as Rachel Carson, people today
are becoming aware of the effects of DDT and now
DDT is banned for use.
8Herbivory
- Herbivory an interaction in which an herbivore
eats parts of plants or alga - DDT is sprayed all over plants for the purpose of
killing unwanted insects. Because DDT is sprayed
on plants, herbivores that eat those plants will
take in the DDT. If the herbivores take in the
DDT in large consumptions, then they will be
unable to carry out necessary life functions,
like reproduction, and eventually become extinct.
Hence there wouldnt be anymore interaction
between the involved plant species and their
herbivore predators. - Global warming also affects herbivory. Global
warming is when the global temperature increases.
Many organisms, especially plants that cannot
disperse rapidly over long distances, would
probably not be able to survive the high rates of
climate change projected to result from global
warming. If plants, dont survive, then neither
will the herbivores, which are dependent on them.
9Predation
- Predation An interaction between species in
which one species, the predator, eats the other,
the prey. - Predation is also affected severely by DDT. After
DDT was released to the public, one of the first
signs that DDT was a serious threat to the
environment was when the population of various
bird species, such as pelicans, ospreys, eagles,
all birds that feed on herbivores that feed on
plants. The accumulation of DDT in the tissues of
these birds interfered with the deposition of
calcium in their eggshells, a trend that may have
already occurred because of environmental
conditions. When these birds tried to incubate
their shells, the weight of the parents broke the
shells of the affected eggs, resulting a decline
of reproduction rates. Basically, this shows how
the interaction between predators (birds) and
their preys (herbivores) leads to the decline in
the population of the predator (and in some cases
even the prey) species. - Predation is affected by acid precipitation. In a
study in Norway, it was found that fish were
dying in ponds and lakes because of acid rain
from pollutants. This would cause the whole lake
and pond ecosystem to suffer not only due to the
acid precipitation, but the loss of one species
would indicate the loss of the predator species
and eventually the elimination of certain food
chains.
10Interspecific Competition
- Interspecific Competition competition for
resources between plants, between animals, or
between decomposers when resources are in short
supply. - DDT affects interspecific competition.
Competition can limit herbivore numbers because
competition may maintain herbivore numbers below
what the vegetation could feed. But as mentioned
earlier herbivore numbers can be easily reduced
through DDTs affect on herbivory. This will
immediately affect competition. The worse part is
that competition includes more than one species,
so if an herbivore species, which was controlled
by the competing species is eliminated, then the
competing species will have nothing to compete
for and one of two things will happen
(1)competition will end and the competing will
find different, new preys or (2) the competing
species will die of starvation. Basically, DDT
has an indirect relationship with competition,
but it still affects the outcome of interspecific
competition. - Competition is affected by the overloaded global
carrying capacity because as we approach or
exceed the carrying capacity the amount of
available resources will not suffice for the
amount of organisms. Hence competition will be
brought to a new level and many species may face
competitive exclusion.