?? ???? Calx ?? ????? ??? ???. ??? ?? ??? 1807 ???? Davy. Knop ?? ??? 1860 ???? ??? ?? ???? ?????? ????? ????? ???.

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• ?? ???? Calx ?? ????? ??? ???. ??? ?? ??? 1807
  ???? Davy. Knop ?? ??? 1860 ???? ??? ?? ????
  ?????? ????? ????? ???.
• ????? ?? ??? ?? ??? ????? ???? ?? ???? ?? ??? ?
  ??? ?? ?? ???? ??? ???? ??? ?? ??? ???? ??? ????
  ????? ?? ???.
• ?? ????? ?? ?? ??? ???? ???? ???? ??? ??? ?????
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• ????????
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• ???? ????
• ????? ?? ???? ???? ?? ???? ????? ???? ????? ??
  ???.
• ???? ?? ?? ????? ????? ????? ????? ???.

4
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• ?? ???? ???? ????? ???? ? ????? ???? ?? ?? ?????
  ????? ?????.

3
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• ??? ?? ? ????? ???? ?? ????
• ????? ???? ?? ???????? ? ?????? ? ?? ????? ????
  ???? ?? ????????? ???? ????.
• ??? ???? ?? ?? ???? ????? ?????? ????? ???.
• ?? ????? ???? ?? ????? ?? ?? ?????? ?????
  ???????? ?????? ? ????.
• ?????? ?????-????????? ???? ????? ???? ????? ??
  ????? ATPase ? NAD-Kinase ??? ? ???? ????? ?????
  ??????? ?????? ?? ????
• ???? ????? ?????? ???? ?? ????? ??? ????
  ?????????
• ??? ???? ????? ???? ?????? ???????????? ? ???????
• ????? ??? ?????? ???? ?????

6
7
????? ? ??????? ???????

• ?? ?????????? ?????? ?? ???? ????? ????? ????? ??
  ???
• Blossom End Rot
• Cracking fruits
• Tipburn in lettuce
• Bitter pit in apples
• ????? ?? ?? ?????? ? ???? ?? ??? ???? ?? ???
  ?????? ??? ?????? ???? ?? ? ??????? ?? ???? ?????
  ?? ?? ?? ?????? ???.

7
8
????? ? ????? ?????? ?????

• ????? ? ??? ?? ?? ?????? ?? ??? ??????
• Poinsettia
• Tulip
• ????? ? ??? ?? ?? ?????? ?? ??? ???? ?????
• Roses
• Gerbera
•    

8
9
Blossom End Rot
5
10
Tip burn
10
11
Bitter Pit
11
12
Cracking fruits
12
13
Poinsettia Bract necrosis
13
14
Poinsettia Bract necrosis
Herbaugh and Woltz, 1989 Herbaugh and Woltz,
1986
14
15
Poinsettia Lateral Stem Breakage
15
16
Tulip
??? ? ?????????? ???? ?? ????
16
17
Tulip
Nelson and Niedziela, 1998 a Nelson and
Niedziela, 1998 b Nelson et al., 2003
17
18
Cut roses
18
19
Cut roses
Ganmore-Neumann and Davidov, 1993
19
20
Cut roses
Torre et al., 2001
20
21
Cut roses
- Ca
Ca
b
a
Torre et al., 1999
21
22
Gerbera
?? ????? ??? ? ?????? ???? ????? ?? ??? ?? ??
????? ??? ?????? ?? ???? ????? ?? ???? ???? ??
???.
22
23
Gerbera
Gerasopoulos and Chebli, 1999
23
24
????? ????

• ????? ????? ?????? ????? ?? ?? ??? ??? ?? ?????
  ??? ????? ???? ???
• ??????? ?????? ?? ?????? ??? ????? ?? ????
  ??????? ???? ?????
• ??? ???? ???? ?????
• ???? ?? ????? ???? ??????? ?????
• ????? ??????/?????? ????? ????????? ? ?????
• ???????? ????? ?? ????? ????? ???? ?? ????? ?????
  ?????? ?? ???? ?? ?? ???? ????? ?????????? ??????
  ??? ?? ?? ???? ?? ???.

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?????

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  influence of humic acids derived from
  earthworm-processed organic wastes on plant
  growth. Bioresearch Tech. 847-14
• Bidegain, R. A. M. Kaemmerer M. Guiresse M.
  Hafidi. 2001. Effect of humic substances from
  composted or chemically decomposed polar sawdust
  on mineral nutrition of ryegrass. J. of Agri.
  Sci. Cambridge. 134 259-267
• Chunhua, L. R. S. Cooper D. C. Bowman. 1998.
  Humic acid application affects photosynthesis,
  root development and nutrition content of
  Creeping Bentgrass. Hortscience, 33(6) 1023-1025
• Cooper, R. J. L. Chunhua D. S. Fisher. 1998.
  Influence of humic substances on rooting and
  nutrient content of creeping bentgrass. Crop
  Science. 38 1639-1644
• Delfine, S R. Tognetti E. Desiberio A.
  alvino. 2005. Effect of foliar application of N
  and humic acids on growth and yield of durum
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• Gerasopoulos, D. B. Chebli. 1999. Effects of
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  vase life of Gerbera. J. of Hort. Sci. and
  Biotech. 74(1) 78-81
• Gislerod, H. R. 1999. The role of calcium on
  several aspects of plant and flower quality from
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  345-352

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1. Hartwigsen, J. A. M. R. Evans.
   2000. Humic acid seed and substrate treatments
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   Ethylene production, ACC content, PAL and POD
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   Biochem. 34 1527-1536
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7. Pilanali, N. M. Kaplan. 2003. Investigation on
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1. Pizzeghello, D G. Nicolini S. Nardi. 2002.
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   attributes and treatment with flower
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   level of calcium in nutrient solution improves
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